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Studies on development of antheridia and antheridial dehiscence have been conducted by Davie and Hartman.
Approximately 50% of the antheridial primordial protoplasts obtained by enzymatic isolation were found to be viable.
Each antheridial chamber incorporates several antheridia, all of which are developed from one antheridial initial.
The latter are much more abundant in the antheridial tufts, where they have somewhat swollen heads.
Fern gametes are free-swimming and a thin film of water is required for the male gametes to travel from the antheridia into the neck of the archcgonia to fertilize the eggs.
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