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The internodes have invaginations that are termed cupules .
Lyginopterids bore cupules , which can be described as tulip shaped structures.
Particle capture was by cilia located in the cupules .
Thus, the cupule is a simple uniaxial structure.
The ovule is surrounded by integument tissues which produce the seed coat, and in the earliest seed plants another layer called the cupule enclosed the entire ovule/seed.
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