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The average number of the repetitions is shown as a subscript index
Here and henceforth, the sum is over all mutations, the subscript E denotes segregating [equilibrium (E)] mutations, and the overbar indicates the arithmetic mean.
Of course we have modernised the notation, for example subscript notation was not used in Parseval's time, and we have also corrected his theorem for he omitted the first 2 on the left hand side.
But, what is interesting here is that this particular industrial strength beast has subscript keys.
We refer to the frequencies of double reduction and recombination fraction between the markers for parent P by, ß, and without the subscript P, unless otherwise specified.
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