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Sometimes grana lost their stacked organization and appeared to slide partially apart at the partitions.
The thylakoid system was normally developed with grana composed of, on average, three to four stacked thylakoids.
The VPC chloroplasts possessed well-developed thylakoids with grana .
The chloroplasts with large grana stacks were seen in 50% of cases but none of the parameters significantly explained the occurrence.
Chloroplast envelopes, stacked grana and unstacked stroma thylakoids were well-defined.
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