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It seems clear that the photocyte has substantial local control of its luminescence and is not dependent on external oxygen in any simple way.
The luminescence in these cases may be excited by direct electron impact rather than with UV light.
Intriguingly, rhythmic luminescence in certain lines was affected by only a subset of the pacemaker mutations.
Pholasin alone can emit luminescence if exposed to superoxide.
The luminescence was quantified in live cells using bioluminescence imaging.