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The applicability of the theory can be improved by considering inhomogeneous densities in all three dimensions.
An explanation for this may lie in the inhomogeneous nature of tumors that often have a necrotic center that can represent up to 50-60% of the tumors' weight.
Among the remainder were embryos that arrested at various stages of development with many having an inhomogeneous density of nuclei.
However, in the cell, molecular behavior is very inhomogeneous : even molecules of single species interact stochastically with distinct molecules or cellular structures in a variety of local environments.
Such inhomogeneous distribution within the membrane would induce an inhomogeneous increase of the lateral pressure profile which, in turn, would specifically affect the activity of membrane proteins.
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